In order to celebrate the birthday of his beloved little brother Rookie (voiced by Christopher Knights), the boss (voiced by Tom McGrath) leads rookie and good partners Koski (voiced by Chris Miller) and Rego into the most heavily defended U.S. Gold Reserve Center, only to find the only vending machine selling French fries at the end of the vault. Who knew that not long after they completed the mission, they were swallowed up by a tentacled vending machine and then transported to a mysterious place. In the fully enclosed facility, four penguins meet Dave the octopus (voiced by John Malkovich) disguised as a genetic scientist. It turns out that Dave was once a star octopus in the zoo, but with the arrival of the cute penguins, he fell into the valley of despair. So Dave immersed himself in the development of the terrible serum, vowing to take revenge on all the penguins in the zoo where he had stayed.
The boss and his friends are rescued by the North Wind Agents on the way to escape, and they and these arrogant Agents will each stop Dave's plot...
These four penguins first appeared in the movie Madagascar. As indispensable supporting roles like the squirrels in "Ice Age", these four penguins have significantly more scenes than squirrels. Unlike Guardians of the Galaxy, which would have been nothing to watch without Raccoons and Treemen, Madagascar would have been like a movie without Penguins and King Julian, but the story is completely different. This may be why DreamWorks later produced three seasons of animated films specifically for Penguins and King Julian.
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